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Type material
Holotype ♂. AUSTRALIA: Western Australia: Cape Range National Park, ca. 6 km NE of Milyering Visitor Centre, site TZ17C, 21.99285°S, 113.97426°E, 24 June 2019, walking, day, 2019-048, M.S. Harvey, N. Tatarnic, A.L. McMah (WAM T148118).
Diagnosis
Males of Missulena ignea sp. nov. are morphologically closest to M. mainae Miglio Harms, Framenau & Harvey, 2014 and M. minima sp. nov., which also have a bright red pars cephalica, pars thoracica, and chelicerae. They differ from these species by the presence of a rasp on patella I–IV in M. ignea sp. nov., whilst only III and IV have a rasp in the other species (Fig. 8k–n vs Fig. 11k–n, fig. 10a, b Miglio et al. 2014). They further differ by the presence of scopulae on tarsus III, IV and metatarsus III, IV in M. ignea sp. nov., which are only on metatarsus IV in these other species, and by by the absence of a swollen metatarsus IV in M. ignea sp. nov. (Fig. 8j vs Fig. 11j, fig. 10k Miglio et al. 2014).
Description
Male (based on holotype; WAM T148118). Total length 8.15.
Colour in ethanol: Pars cephalica, pars thoracica, and chelicerae reddish-orange (Fig. 8a); black pigmentation around AME, PLE, and ALE; PME pale (Fig. 8a, e). Sternum pale orange, with eight sigilla of similar colour (Fig. 8b); labium and maxillae darker orange with faint brown latitudinal striations on the base of labium (Fig. 8b, f). Abdomen with a strong, metallic blue-grey sheen dorsally, ventrally black; spinnerets pale violet grey, with white interconnecting segments (Fig. 8c, d). Legs dark olive-brown, femur dorsal surfaces with a faint blue metallic sheen, more prominent on legs I, II; tarsi of all segments paler brown, blending to pale cream apically.
Carapace: 3.91 long and 4.57 wide; pars cephalica height 1.62; pars cephalica covers 0.58 of carapace length. Pars cephalica with moderate granulation and with scattered setae. Pars thoracica granulation less coarse than that of pars cephalica, with two lateral indentations posteriorly, and a deep central posterior notch; lateral edges strongly recurved and undulating (Fig. 8a, i). Pars cephalica moderately elevated (Fig. 8i). Eyes: OQ 3.52 times wider than long; anterior eyes straight; posterior eyes strongly recurved (Fig. 8e). Chelicerae: 2.72 long; edges rounded and broadly cylindrical; dorsal surface smooth, granulations and regular longitudinal ridges on prolateral surfaces; faint ridges on retrolateral surfaces (Fig. 8a). Sternum: ovoid; with unevenly scattered setae, denser along margins and anterior. Four pairs of sigilla, all lightly depressed, anterior pair very small, indistinct and oval; second pair small, circular; third pair ca. three times larger than the second pair and oval; fourth pair largest, ca. three times larger than the third pair, oval (Fig. 8a). Labiosternal junction defined, fused (Fig. 8a). Cheliceral furrow with row of eight teeth prolaterally (seven right), five teeth retrolaterally (seven right) and with a cluster of five small teeth in the inter-furrow region (Fig. 8h). Maxillae: with covering of setae, denser anteriorly, and with ca. 50 cuspules on anterior and prolateral margins (Fig. 8f). Labium: squat pyramidal, with sparse long setae and with ca. 17 cuspules anteriorly (Fig. 8f). Rastellum present, not well pronounced, with five (right six) strong, conical spines (Fig. 8g). With ca. 20 setae forming a fringe over the base the fang of each chelicera - the setae being thicker retrolaterally.
Abdomen: 4.02 long and 3.37 wide; oval; four spinnerets (Fig. 8c, d).
Legs: scattered brown-black setae on all surfaces. Leg spination: leg I: tibia rv3, v9, pl0, d0; metatarsus rv6, v5, pl0, d0; tarsus rv2, v0, pl0, d0; leg II: tibia rv6, v6, pl0, d0; metatarsus rv7, v2, pl4, d0; tarsus rv2, v0, pl0, d0; leg III: tibia rv1, v3, pl3, d4; metatarsus rv6, v0, pl3, d7; tarsus rv5, v3, pl4, d2; leg IV: tibia rv2, v5, pl0, d0; metatarsus rv8, v0, pl4, d2; tarsus rv8, v3, pl9, d4. Patella I with six spines prolaterally, four ventral spines, one spine retrolaterally (Fig. 8k); patella II with two prolateral spines apically, two ventral spines apically, and one retrolateral spine apically (Fig. 8l); patella III with 18 spines on the prolateral and dorsal surfaces (Fig. 8m); patella IV with ca. six spinules dorsally interspersed amongst elongate setae (Fig. 8n). Scopulae on ventral surfaces of tarsi and metatarsi III and IV (Fig. 8j, leg IV imaged). Claws long. Leg measurements: Leg I: femur 3.05, patella 1.90, tibia 2.26, metatarsus 1.97, tarsus 1.42, total 10.06. Leg II: femur 2.37, patella 1.74, tibia 1.86, metatarsus 1.92, tarsus 1.24, total 9.13. Leg III: femur 1.95, patella 1.52, tibia 1.40, metatarsus 1.99, tarsus 1.49, total 8.35. Leg IV: femur 2.75, patella 1.97, tibia 2.41, metatarsus 2.04, tarsus 1.62, total 10.79. Formula: 4>1>3>2.
Pedipalp: femur 2.55, patella 1.52, tibia 2.28, tarsus 0.69; all segments with scattered setae, those of the tibia thicker and longer ventrally (Fig. 9a–c). Tibia thin, broader basally, 1.45 wide on the widest point from dorsal and prolateral view (Fig. 9a, b). Bulb roughly pyriform (Fig. 9a, b). Embolus short, small swelling near base; tip squared, without an embolar tooth (Fig. 9d).
Variation. Only known from holotype male.
Etymology
The specific epithet is a Latin adjective igneus meaning ‘fiery’, and refers to the bright orange-red colouration of the pars cephalica, pars thoracica and chelicerae of this species.
Distribution
Only known from Cape Range National Park, mid-western WA (Fig. 10).